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Four-year PhD Studentship Programmes

Red blood cells
Our flagship scheme that supports the most promising students to undertake postgraduate training.
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Purpose

This is a flagship scheme aimed at supporting the most promising students to undertake in depth postgraduate training.

We currently support 32 Programmes based in centres of excellence throughout the UK with specialised training provided in a range of important biomedical research areas:

  • developmental biology and cell biology
  • genetics, statistics and epidemiology
  • immunology and infectious disease
  • molecular and cellular biology
  • neuroscience
  • physiological sciences
  • structural biology and bioinformatics.

In addition, we also fund a Four-year PhD Programme at the Wellcome Trust Sanger Institute. Please contact the Institute directly for further information.

The first year of a Programme combines taught courses with laboratory rotations to broaden the student's knowledge of the subject area.

At the end of the first year, students will make an informed choice of their three-year PhD research project.

Note: if you are looking for the Trust’s recent call for four-year PhD programmes, please visit this page.

Programmes

For information on the 2016 uptake, please contact the following programmes directly:

Developmental biology and cell biology

Developmental Mechanisms, University of Cambridge
Director: Professor Daniel St Johnston

Stem Cell Biology and Medicine, University of Cambridge
Director: Professor Austin Smith

Chromosome and Developmental Biology, University of Oxford
Director: Professor Elizabeth Robertson

Cell Therapies and Regenerative Medicine, King’s College London
Director: Professor Fiona Watt

Genetics and population health

Mathematical Genomics and Medicine, University of Cambridge
Director: Professor Simon Tavaré

Molecular, Genetic and Lifecourse Epidemiology, University of Bristol
Director: Professor George Davey-Smith

Genomic Medicine and Statistics, University of Oxford
Directors: Professor Julian Knight

Epidemiology, Evolution and Control of Infectious Disease, Imperial College London
Director: Professor Azra Ghani

Public Health Economics and Decision Science, University of Sheffield
Director: Professor Petra Meier

Immunology and infectious disease

Infection, Immunity and Inflammation, University of Cambridge
Director: Professor Paul Lehner

Molecular and Cellular Basis of Infection, Imperial College London
Directors: Professor Murray Selkirk and Professor Martin Allday

Immunology, Infection and Translational Medicine, University of Oxford
Director: Professor Fiona Powrie

Antimicrobials and Antimicrobial Resistance, University of Nottingham and University of Birmingham
Directors: Professor Paul Williams and Professor Ian Henderson

Mechanisms of Inflammatory Disease, University of Birmingham
Director: Steve Watson

Hosts, Pathogens and Global Health, University of Edinburgh
Director: Professor Judith Allen

Molecular and cellular biology

Dynamic Cell Biology, University of Bristol
Director: Professor Peter Cullen

Molecular and Cellular Biology, University of Dundee
Director: Professor Robert Ryan

The Dynamics of Cellular Regulatory Networks, University of Manchester
Director: Professor Hilary Ashe

Quantitative and Biophysical Biology, University of Manchester
Director: Professor Christopher Thompson

Neuroscience

Neural Dynamics: from synapses to systems in health and disease, University of Bristol
Director: Professor Richard Apps

Integrative Neuroscience, University of Cardiff
Director: Professor John Aggleton

Neuroscience, University of Oxford
Director: Professor Andrew King

Neuroscience, University College London
Director: Professor David Attwell

OXION-Membrane Transport and Disease
Director: Dr Stephen Tucker

Translational Neuroscience – life-course influences on human brain health, University of Edinburgh
Director: Professor Charles ffrench-Constant

Physiological sciences

Metabolic and Cardiovascular Disease, University of Cambridge
Director: Professor Stephen O'Rahilly

Cellular and Molecular Physiology, University of Liverpool
Director: Professor Ian Prior

Tissue Repair, University of Edinburgh
Director: Professor Stuart Forbes

Structural biology and bioinformatics

Interdisciplinary Programme in Structural, Computational and Chemical Biology, University College London, Birkbeck College and The Francis Crick Institute
Director: Professor Gabriel Waksman

Molecular Basis of Biological Mechanisms, University of Leeds
Director: Professor Alan Berry

Cellular Structural Biology, University of Oxford
Director: Professor David Stuart

Theoretical Systems Biology and Bioinformatics, Imperial College London
Director: Professor Michael Sternburg

Active programmes that are not recruiting

Developmental and Stem Cell Biology, University College London
Director: Professor Claudio Stern

Systems Neuroscience: from networks to behaviour, University of Newcastle Upon Tyne
Director: Stuart Baker

Cell Biology, University of Edinburgh
Director: Professor Jean Beggs

Computational Infection Biology, University College Dublin
Director: Professor Geraldine Butler

Combating infectious disease: computational approaches in translational science (CIDCATS), University of York
Director: Professor Paul Kaye

What's included

Support is provided for four years and includes:

  • a stipend
  • PhD registration fees at UK/EU student rate
  • contribution towards laboratory rotation expenses in the first year
  • research expenses for years two to four
  • contribution towards travel
  • contribution towards transferable-skills training.

Eligibility

You should be a student who has, or expects to obtain, a first- or upper-second-class honours degree or equivalent.

Application process

Students are recruited annually by the individual Programmes for uptake in October each year. Recruitment begins in the preceding December.

If you are interested in applying you should contact the relevant Programme directly. Please see their websites for more details (listed under the ‘Programmes’ tab).

Deadlines

Students are recruited annually by the individual Programmes for uptake in October each year. Recruitment begins in the preceding December.

Contacts

Enquiries should be directed to relevant Programme. Please see their websites for more details (listed under the ‘Programmes’ tab).

Please email any policy enquiries to 4yrphd@wellcome.ac.uk.

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